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484 DIGESTION.
and Cohnheim.1
These investigations refer to the conditions in dogs,
and as shown by Rosenfeld 2
in horses, and by Lotsch 3 in pigs, that
the conditions are different in other animals. The following description
applies only to dogs.
In the dog Abderhalden, London and co-workers 4
have shown
that in the stomach proteoses and peptones are formed, but no amino-
acids, or at least not in any mentionable quantity. The scanty occurrence
-of amino-acids is substantiated by the observations of Zunz and others 5
that the amount of amino-nitrogen titratable with formol in the stomach
contents, is only small.
In like manner we must agree in the belief that a part of the protein
always leaves the stomach undigested and that the principal mass, about
80 per cent, passes into the intestine more or less digested. Besides this
it also seems as if the peptones occur in the pylorus part to a greater
extent than the proteoses, while in the fundus part the reverse is the
case. Of the dissolved protein of the entire stomach-contents about
60 per cent exists as proteoses. Opinions are also contradictory in
regard to the absorption of the decomposition products of the proteins
in the stomach. While several investigators, like Tobler, Lang, Cohn-
heim, Zunz and others accept such an absorption, London and co-workers
positively deny this.
The digestion of sundry foods is not dependent on one organ alone,
but is divided among several. For this reason it is to be expected that
the various digestive organs can act for one another to a certain extent,
and that therefore the work of the stomach could be taken up more or
less by the intestine. This in fact is the case. Thus the stomachs of
dogs and cats have been completely extirpated or nearly so (Czerny,
Carvallo and Pachon, London and collaborators), or that part
necessary in the digestive process has also been eliminated by plugging
the pyloric opening (Ludwig and Ogata), and in both cases it was pos-
sible to keep the animal alive, well fed, and strong for a shorter or
longer time. The extirpation of the stomach has also been repeatedly
1
Tobler, Zeitschr. f. physiol. Chem., 45; Lang, Bioch. Zeitschr., 2; Cohnheim,
Munch, med. Wochenschr., 1907. In regard to the works of Zunz, London, and
collaborators, see footnotes 1, 2 and 3, p. 482.
2
Rosenfeld, Ueber die Eiweissverdauung im Magen des Pferdes, Inaug.-Dissert.,
Dresden, 1908.
’ Lotsch. Zur Kenntnis der Verdauung von Fleisch im Magen und Dunndarm des
Schweines, Inaug.-Dissert. Freiburg i. Sa., 1908; see also Abderhalden, Klingemann
and Pappenhusen, Zeitschr. f. physiol. Chem. 71, 411 (1911).
4
Abderhalden and London, with Kautsch, Zeitschr. f. physiol. Chem., 48, with
L. Baurnann, ibid., 51, and with v. Korosy, ibid., 51.
:
Intern. Beitr. zu Path. u. Ther. d. Ern.-Stor. 2; London and Rabinowitsch,
Zeitschr. f. physiol. Chem. 74.
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